The order of presentation of virus orders, families and genera does not reflect any hierarchical or phylogenetic classification, but only a convenient order of presentation. Since a taxonomic structure above the level of family or genus has not been developed systematically (with the exception of three orders Caudovirales, Mononegavirales and Nidovirales) any sequence of listing of taxa must be arbitrary. The order of presentation of the virus families and genera of the VIIth ICTV Report follows four criteria: (1) the nature of the viral genome, (2) standedness of the viral genome, (3) the fact that some viruses are reverse transcribed, and (4) the polarity of the virus genome. As there are no known ssDNA, nor dsRNA reverse transcribed viruses, and there are negative sense viruses only for ssRNA viruses, these four criteria give rise to seven clusters comprising the 56 families and 233 genera of viruses. In addition, subviral agents, namely the satellites, viroids and agents of spongiform encephalopathies (prions) are included, in most cases official taxonomic status. Finally, a list of unassigned viruses is provided.

¹ Quotes are used to denote taxon names that are not ICTV International approved names, and are thus temporary until formal names are approved.

² “hosts also include plants (genus Pseudovirus)”

³ “hosts also include fungi (genus Hemivirus)”

⁴ “hosts also include fungi, plants and invertebrates (genus Metavirus)”

^* Vertebrate or plant arthropod-borne viruses are listed according to their vertebrate or plant hosts and are identified by an asterisk.


Order
Family	Subfamily	Genus	Type species	Host	Page

The DNA Viruses
The dsDNA Viruses

*CAUDOVIRALES*
*Myoviridae*
		“T4-like viruses”¹	Enterobacteria phage T4	Bacteria
		“P1-like viruses”	Enterobacteria phage P1	Bacteria
		“P2-like viruses”	Enterobacteria phage P2	Bacteria
		“Mu-like viruses”	Enterobacteria phage Mu	Bacteria
		“SP01-like viruses”	Bacillus phage SP01	Bacteria
		“H-like viruses”	Halobacterium virus H	Archaea
Siphoviridae		“-like viruses”	Enterobacteria phage	Bacteria
		“T1-like viruses”	Enterobacteria phage T1	Bacteria
		“T5-like viruses”	Enterobacteria phage T5	Bacteria
		“L5-like viruses”	Mycobacterium phage L5	Bacteria
		“c2-like viruses”	Lactococcus phage c2	Bacteria
		“M1-like viruses”	Methanobacterium virus M1	Archaea
Podoviridae		“T7-like viruses”	Enterobacteria phage T7	Bacteria
		“P22-like viruses”	Enterobacteria phage P22	Bacteria
		“29-like viruses”	Bacillus phage 29	Bacteria

Tectiviridae		Tectivirus	Enterobacteria phage PRD1	Bacteria

Corticoviridae		Corticovirus	Alteromonas phage PM2	Bacteria

Plasmaviridae		Plasmavirus	Acholeplasma phage L2	Mycoplasma

Lipothrixviridae		Lipothrixvirus	Thermoproteus virus 1	Archaea

Rudiviridae		Rudivirus	Sulfolobus virus SIRV1	Archaea

Fuselloviridae		Fusellovirus	Sulfolobus virus SSV1	Archaea

		“Sulfolobus SNDV-like viruses”	Sulfolobus virus SNDV	Archaea

Poxviridae

	Chordopoxvirinae	Orthopoxvirus	Vaccinia virus	Vertebrates
		Parapoxvirus	Orf virus	Vertebrates
		Avipoxvirus	Fowlpox virus	Vertebrates
		Capripoxvirus	Sheeppox virus	Vertebrates
		Leporipoxvirus	Myxoma virus	Vertebrates
		Suipoxvirus	Swinepox virus	Vertebrates
		Molluscipoxvirus	Molluscum contagiosum virus	Vertebrates
		Yatapoxvirus	Yaba monkey tumor virus	Vertebrates
	Entomopoxvirinae	Entomopoxvirus A	Melolontha melolontha entomopoxvirus	Invertebrates
		Entomopoxvirus B	Amsacta moorei entomopoxvirus	Invertebrates
		Entomopoxvirus C	Chironomus luridus entomopoxvirus	Invertebrates
Asfarviridae		Asfivirus	African swine fever virus	Vertebrates^*

Iridoviridae
		Iridovirus	Invertebrate iridescent virus 6	Invertebrates
		Chloriridovirus	Invertebrate iridescent virus 3	Invertebrates
		Ranavirus	Frog virus 3	Vertebrates
		Lymphocystivirus	Lymphocystis disease virus 1	Vertebrates

Phycodnaviridae
		Chlorovirus	Paramecium bursaria Chlorella virus 1	Algae
		Prasinovirus	Micromonas pusilla virus SP1	Algae
		Prymnesiovirus	Chrysochromulina brevifilum virus PW1	Algae
		Phaeovirus	Ectocarpus siliculosus virus 1	Algae

Baculoviridae
		Nucleopolyhedrovirus	Autographa californica nucleopolyhedrovirus	Invertebrates
		Granulovirus	Cydia pomonella granulovirus	Invertebrates

Herpesviridae
	Alphaherpesvirinae
		Simplexvirus	Human herpesvirus 1	Vertebrates
		Varicellovirus	Human herpesvirus 3	Vertebrates
		“Marek’s disease-like viruses”	Gallid herpesvirus 2	Vertebrates
		“Infectious laryngotracheitis-like viruses”	Gallid herpesvirus 1	Vertebrates
	Betaherpesvirinae
		Cytomegalovirus	Human herpesvirus 5	Vertebrates
		Muromegalovirus	Murid herpesvirus 1	Vertebrates
		Roseolovirus	Human herpesvirus 6	Vertebrates
	Gammaherpesvirinae
		Lymphocryptovirus	Human herpesvirus 4	Vertebrates
		Rhadinovirus	Saimiriine herpesvirus 2	Vertebrates

		“Ictalurid herpes-like viruses”	Ictalurid herpesvirus 1	Vertebrates

Adenoviridae
		Mastadenovirus	Human adenovirus C	Vertebrates
		Aviadenovirus	Fowl adenovirus A	Vertebrates

		Rhizidiovirus	Rhizidiomyces virus	Fungi

Polyomaviridae		Polyomavirus	Simian virus 40	Vertebrates

Papillomaviridae		Papillomavirus	Cottontail rabbit papillomavirus	Vertebrates

Polydnaviridae
		Ichnovirus	Campoletis sonorensis ichnovirus	Invertebrates
		Bracovirus	Cotesia melanoscela brachovirus	Invertebrates

Ascoviridae		Ascovirus	Spodoptera frugiperda ascovirus	Invertebrates
The ssDNA Viruses

Inoviridae
		Inovirus	Enterobacteria phage M13	Bacteria
		Plectrovirus	Acholeplasma phage MV-L51	Mycoplasma
Microviridae
		Microvirus	Enterobacteria phage X174	Bacteria
		Spiromicrovirus	Spiroplasma phage 4	Spiroplasma
		Bdellomicrovirus	Bdellovibrio phage MAC1	Bacteria
		Chlamydiamicrovirus	Chlamydia phage 1	Bacteria
Geminiviridae
		Mastrevirus	Maize streak virus	Plants
		Curtovirus	Beet curly top virus	Plants
		Begomovirus	Bean golden mosaic virus-Puerto Rico	Plants
Circoviridae		Circovirus	Chicken anemia virus	Vertebrates

		Nanovirus	Subterranean clover stunt virus	Plants

Parvoviridae
	Parvovirinae
		Parvovirus	Mice minute virus	Vertebrates
		Erythrovirus	B19 virus	Vertebrates
		Dependovirus	Adeno-associated virus 2	Vertebrates
	Densovirinae
		Densovirus	Junonia coenia densovirus	Invertebrates
		Iteravirus	Bombyx mori densovirus	Invertebrates
		Brevidensovirus	Aedes aegypti densovirus	Invertebrates
The DNA and RNA Reverse Transcribing Viruses

Hepadnaviridae
		Orthohepadnavirus	Hepatitis B virus	Vertebrates
		Avihepadnavirus	Duck hepatitis B virus	Vertebrates

Caulimoviridae
		Caulimovirus	Cauliflower mosaic virus	Plants
		“Petunia vein-clearing-like viruses”	Petunia vein-clearing virus	Plants
		“Soybean chlorotic mottle-like viruses”	Soybean chlorotic mottle virus	Plants
		“Cassava vein mosaic virus-like viruses”	Cassava vein mosaic virus	Plants
		Badnavirus	Commelina yellow mottle virus	Plants
		“Rice tungro bacilliform-like viruses”	Rice tungro bacilliform virus	Plants

Pseudoviridae
		Pseudovirus	Saccharomyces cerevisiae Ty1 virus	Fungi²
		Hemivirus	Drosophila melanogaster copia virus	Invertebrates³

Metaviridae
		Metavirus	Saccharomyces cerevisiae Ty3 virus	Fungi⁴
		Errantivirus	Drosophila melanogaster gypsy virus	Invertebrates

Retroviridae
		Alpharetrovirus	Avian leukosis virus	Vertebrates
		Betaretrovirus	Mouse mammary tumor virus	Vertebrates
		Gammaretrovirus	Murine leukemia virus	Vertebrates
		Deltaretrovirus	Bovine leukemia virus	Vertebrates
		Epsilonretrovirus	Walleye dermal sarcoma virus	Vertebrates
		Lentivirus	Human immunodeficiency virus 1	Vertebrates
		Spumavirus	Chimpanzee foamy virus	Vertebrates
The RNA Viruses

The dsRNA Viruses

Cystoviridae		Cystovirus	Pseudomonas phage 6	Bacteria

Reoviridae
		Orthoreovirus	Mammalian orthoreovirus	Vertebrates
		Orbivirus	Bluetongue virus	Vertebrates^*
		Rotavirus	Rotavirus A	Vertebrates
		Coltivirus	Colorado tick fever virus	Vertebrates^*
		Aquareovirus	Aquareovirus A	Vertebrates^*
		Cypovirus	Cypovirus 1	Invertebrates
		Fijivirus	Fiji disease virus	Plants^*
		Phytoreovirus	Rice dwarf virus	Plants^*
		Oryzavirus	Rice ragged stunt virus	Plants^*

Birnaviridae
		Aquabirnavirus	Infectious pancreatic necrosis virus	Vertebrates^*
		Avibirnavirus	Infectious bursal disease virus	Vertebrates
		Entomobirnavirus	Drosophila X virus	Invertebrates

Totiviridae
		Totivirus	Saccharomyces cerevisiae virus L-A	Fungi
		Giardiavirus	Giardia lamblia virus	Protozoa
		Leishmaniavirus	Leishmania RNA virus 1-1	Protozoa

Partitiviridae
		Partitivirus	Gaeumannomyces graminis virus 019/6-A	Fungi
		Chrysovirus	Penicillium chrysogenum virus	Fungi
		Alphacryptovirus	White clover cryptic virus 1	Plants
		Betacryptovirus	White clover cryptic virus 2	Plants

Hypoviridae		Hypovirus	Cryphonectria hypovirus 1-EP713	Fungi

		Varicosavirus	Lettuce big-vein virus	Plants
The Negative Sense ssRNA Viruses

MONONEGAVIRALES
Bornaviridae
		Bornavirus	Borna disease virus	Vertebrates
Filoviridae
		“Marburg-like viruses”	Marburg virus	Vertebrates
		“Ebola-like viruses”	Zaire Ebola virus	Vertebrates
Paramyxoviridae
	Paramyxovirinae
		Respirovirus	Sendai virus	Vertebrates
		Rubulavirus	Mumps virus	Vertebrates
		Morbillivirus	Measles virus	Vertebrates
	Pneumovirinae
		Pneumovirus	Human respiratory syncytial virus	Vertebrates
		Metapneumovirus	Turkey rhinotracheitis virus	Vertebrates
Rhabdoviridae
		Vesiculovirus	Vesicular stomatitis Indiana virus	Vertebrates^*
		Lyssavirus	Rabies virus	Vertebrates
		Ephemerovirus	Bovine ephemeral fever virus	Vertebrates^*
		Novirhabdovirus	Infectious hematopoietic necrosis virus	Vertebrates
		Cytorhabdovirus	Lettuce necrotic yellows virus	Plants^*
		Nucleorhabdovirus	Potato yellow dwarf virus	Plants^*

Orthomyxoviridae
		Influenzavirus A	Influenza A virus	Vertebrates
		Influenzavirus B	Influenza B virus	Vertebrates
		Influenzavirus C	Influenza C virus	Vertebrates
		Thogotovirus	Thogoto virus	Vertebrates^*

Bunyaviridae
		Bunyavirus	Bunyamwera virus	Vertebrates^*
		Hantavirus	Hantaan virus	Vertebrates
		Nairovirus	Dugbe virus	Vertebrates^*
		Phlebovirus	Rift Valley fever virus	Vertebrates^*
		Tospovirus	Tomato spotted wilt virus	Plants^*

		Tenuivirus	Rice stripe virus	Plants^*

		Ophiovirus	Citrus psorosis virus	Plants

Arenaviridae		Arenavirus	Lymphocytic choriomeningitis virus	Vertebrates

		Deltavirus	Hepatitis delta virus	Vertebrates

The Positive Sense ssRNA Viruses
Leviviridae
		Levivirus	Enterobacteria phage MS2	Bacteria
		Allolevivirus	Enterobacteria phage Q	Bacteria

Narnaviridae
		Narnavirus	Saccharomyces cerevisiae narnavirus 20S	Fungi
		Mitovirus	Cryphonectria parasitica mitovirus 1-NB631	Fungi

Picornaviridae
		Enterovirus	Poliovirus	Vertebrates
		Rhinovirus	Human rhinovirus A	Vertebrates
		Cardiovirus	Encephalomyocarditis virus	Vertebrates^*
		Aphthovirus	Foot-and-mouth disease virus	Vertebrates
		Hepatovirus	Hepatitis A virus	Vertebrates
		Parechovirus	Human parechovirus	Vertebrates

		“Cricket paralysis-like viruses”	Cricket paralysis virus	Invertebrates

Sequiviridae
		Sequivirus	Parsnip yellow fleck virus	Plants
		Waïkavirus	Rice tungro spherical virus	Plants

Comoviridae
		Comovirus	Cowpea mosaic virus	Plants
		Fabavirus	Broad bean wilt virus 1	Plants
		Nepovirus	Tobacco ringspot virus	Plants

Potyviridae
		Potyvirus	Potato virus Y	Plants
		Ipomovirus	Sweet potato mild mottle virus	Plants
		Macluravirus	Maclura mosaic virus	Plants
		Rymovirus	Ryegrass mosaic virus	Plants
		Tritimovirus	Wheat streak mosaic virus	Plants
		Bymovirus	Barley yellow mosaic virus	Plants

Caliciviridae
		Lagovirus	Rabbit hemorrhagic disease virus	Vertebrates
		“Norwalk-like viruses”	Norwalk virus	Vertebrates
		“Sapporo-like viruses”	Sapporo virus	Vertebrates
		Vesivirus	Swine vesicular exanthema virus	Vertebrates

		“Hepatitis E -like viruses”	Hepatitis E virus	Vertebrates

Astroviridae		Astrovirus	Human astrovirus 1	Vertebrates

Nodaviridae
		Alphanodavirus	Nodamura virus	Invertebrates
		Betanodavirus	Striped jack nervous necrosis virus	Vertebrates*

Tetraviridae
		Betatetravirus	Nudaurelia capensis virus	Invertebrates
		Omegatetravirus	Nudaurelia capensis virus	Invertebrates

		Sobemovirus	Southern bean mosaic virus	Plants

		Marafivirus	Maize rayado fino virus	Plants

Luteoviridae
		Luteovirus	Barley yellow dwarf virus - PAV	Plants
		Polerovirus	Potato leafroll virus	Plants
		Enamovirus	Pea enation mosaic virus-1	Plants

		Umbravirus	Carrot mottle virus	Plants

Tombusviridae
		Aureusvirus	Pothos latent virus	Plants
		Avenavirus	Oat chlorotic stunt virus	Plants
		Carmovirus	Carnation mottle virus	Plants
		Dianthovirus	Carnation ringspot virus	Plants
		Machlomovirus	Maize chlorotic mottle virus	Plants
		Necrovirus	Tobacco necrosis virus	Plants
		Panicovirus	Panicum mosaic virus	Plants
		Tombusvirus	Tomato bushy stunt virus	Plants

Nidovirales
Coronaviridae
		Coronavirus	Infectious bronchitis virus	Vertebrates
		Torovirus	Equine torovirus	Vertebrates
Arteriviridae
		Arterivirus	Equine arteritis virus	Vertebrates

Flaviviridae
		Flavivirus	Yellow fever virus	Vertebrates^*
		Pestivirus	Bovine viral diarrhea virus 1	Vertebrates
		Hepacivirus	Hepatitis C virus	Vertebrates

Togaviridae
		Alphavirus	Sindbis virus	Vertebrates^*
		Rubivirus	Rubella virus	Vertebrates

		Tobamovirus	Tobacco mosaic virus	Plants

		Tobravirus	Tobacco rattle virus	Plants

		Hordeivirus	Barley stripe mosaic virus	Plants

		Furovirus	Soil-borne wheat mosaic virus	Plants

		Pomovirus	Potato mop-top virus	Plants

		Pecluvirus	Peanut clump virus	Plants

		Benyvirus	Beet necrotic yellow vein virus	Plants

Bromoviridae
		Alfamovirus	Alfalfa mosaic virus	Plants
		Bromovirus	Brome mosaic virus	Plants
		Cucumovirus	Cucumber mosaic virus	Plants
		Ilarvirus	Tobacco streak virus	Plants
		Oleavirus	Olive latent virus 2	Plants

		Ourmiavirus	Ourmia melon virus	Plants

		Idaeovirus	Raspberry bushy dwarf virus	Plants

Closteroviridae
		Closterovirus	Beet yellows virus	Plants
		Crinivirus	Lettuce infectious yellows virus	Plants

		Capillovirus	Apple stem grooving virus	Plants

		Trichovirus	Apple chlorotic leaf spot virus	Plants

		Vitivirus	Grapevine virus A	Plants

		Tymovirus	Turnip yellow mosaic virus	Plants

		Carlavirus	Carnation latent virus	Plants

		Potexvirus	Potato virus X	Plants

		Allexivirus	Shallot virus X	Plants

		Foveavirus	Apple stem pitting virus	Plants

Barnaviridae		Barnavirus	Mushroom bacilliform virus	Fungi

Unassigned Viruses	Host	Page

	Vertebrates
	Invertebrates
	Plants
	Fungi

Viroids

Pospiviroidae
	Pospiviroid	Potato spindle tuber viroid	Plants
	Hostuviroid	Hop stunt viroid	Plants
	Cocadviroid	Coconut cadang-cadang viroid	Plants
	Apscaviroid	Apple scar skin viroid	Plants
	Coleviroid	Coleus blumei viroid 1	Plants
Avsunviroidae
	Avsunviroid	Avocado sunblotch viroid	Plants
	Pelamoviroid		Peach latent mosaic viroid	Plants

Subviral Agents: Satellites, and Agents of Spongiform Encephalopathies (Prions)

Subviral Agents	Host	Page
Satellites	Invertebrates
	Plants
	Fungi
Prions	Vertebrates
	Fungi